For me, it's when a dominant character acts submissive, or when a submissive character acts dominant.
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Like when i flirt with Mokou and she gets all mad, or Tewi and I play a prank on Reisen.
@JimT: Oh, oh! I don't know if you remember Myougi Marika from Princess Evangile (stereotypical student council president), but she turns out to be a total sub in the fandisc.
busty subs